Adams, M. A. and P. M. Attiwill (1982). "Nitrogen mineralization and

nitrate reduction in forests." Soil Biol. Biochem. 14(3): 197-202.

The mineralization of soil nitrogen was studied in four forests growing on

krasnozem soils. Soils from Silver Wattle (Acacia dealbata Link.) and

Mountain Ash (Eucalyptus regnans F. Muell.) forests showed considerable

nitrification in laboratory incubations. Messmate (Eucalyptus obliqua

L'Herit) and Monterey Pine (Pinus radiata D. Don) forest soils were

predominantly ammonifiers. Forests having significant soil nitrification

were found to have considerable nitrate reductase activity (NRA) in root or

leaf tissue or both. NRA may therefore be useful as an indication of soil

nitrification in natural ecosystems. The occurrence of nitrification in

Australian forests appears to be predominantly related to the amount of N

present and its rate of turnover rather than to inhibitory effects.

 

Adams, M. A. and P. M. Attiwill (1984). "Role of Acacia Spp. in nutrient

balance and cycling in regenerating Eucalyptus regnans F. Muell. Forests.

I: Temporal changes in biomass and nutrient content." Aust. J. Bot. 32(2):

205-215.

Following severe fire in high-rainfall Eucalyptus regnas F. Muell.

forests, Several Acacia species may germinate in large numbers. Large

amounts of nitrogen as well as calcium, magnesium and potassium are

immobilized in the Acacia biomass, much of which is retunrned to the soil

after canopy closure. Within 3 years of a "regeneration burn", 280 kg N ha

super(-1) was in the above-ground Acacia biomass and litter layer. The

relative abundance of nitrogen in Acacia spp. is a result of N fixation.

The probable rate of N fixation appears significant in relation to losses

of nitrogen associated with slash-burning. It is concluded that the Acacia

spp. may be important in the secondary succession through nutrient

conservation, replacement and redistribution.

 

Adams, M. A. and P. M. Attiwill (1984). "Patterns of nitrogen

mineralization in 23-year old pine forest following nitrogen fertilizing."

For. Ecol. Manage. 7(4): 241-248.

Application of nitrogen fertilizer to 23-year old Pinus radiata D. Don at

high rates induced considerable change in patterns of nitrogen

mineralization in the soil. Increased net mineralization, nitrification,

root Nitrate Reductase Activity, and foliar N levels were measured within

12 months of fertilizer application. Fertilizing did not, however, affect

growth significantly over the 30 months of measurement. Significant

differences in inorganic-N and patterns of mineralization between

fertilized plots and controls were not maintained and all of the fertilized

plots had levels of inorganic-N and patterns of mineralization approaching

those of the control within 30 months. N added to these young aggrading

forests of high productivity therefore appears to be quickly immobilized.

Whithin normal forest rotations, it is apparent that the C/N ratio of the

soil acts as a strong buffer to perturbation of mineralization patterns.

 

Adams, M. A., P. J. Polglase, et al. (1989). "In situ studies of nitrogen

mineralization and uptake in forest soils: Some comments on methodology."

Soil Biol. Biochem. 21(3): 423-429.

Aspects of the methodology of and interpretation of results from, in situ

studies of N-mineralization are discussed with reference to data collected

from 17 eucalypt forests in south-eastern Australia during a 5-year period.

Results suggest that: (i) it is possible to maintain moisture of soils

contained within corers at levels not significantly different from those of

the surrounding soil; (ii) inorganic-N is not produced linearly over time

under field conditions, nor should we expect it to be; (iii) mineralization

rates are affected by all in situ methods. In each of the forests examined,

the average rate of net N-mineralization decreased as the period of

containment increased.

 

Adams, M., P. Attiwill, et al. (1989). "Availability of nitrogen and

phosphorus in forest soils in northeastern Tasmania." Biology and Fertility

of Soils 8(3): 212-218.

Rates of N mineralization and of N uptake were measured in situ in three

eucalypt forests and a cool-temperate rainforest, and were correlated with

productivity. All of the soils had a high capacity for immobilization, and

nitrification was insignificant. Changes in both organic and inorganic P

fractions during in situ containment of soils were small. While the

concentration of inorganic available P was not related to forest

productivity, a measure of labile organic P was closely related both to

productivity and to P in the microbial biomass. Estimates of inorganic- and

organic-N availability were highly correlated with independent estimates of

organic-P availability, and the results are discussed in relation to

biological control of nutrient availability in the surface horizons of

forest soils.

 

Adams, M. and P. Attiwill (1991). "Nutrient balance in forests of northern

Tasmania. 2. Alteration of nutrient availability and soil-water chemistry

as a result of logging, slash-burning and fertilizer application." Forest

Ecology and Management 44(2-4): 115-131.

A 100-ha plot in dry sclerophyll forest in NE Tasmania (dominant species

Eucalyptus obliqua) was logged during summer 1983-84, slash burnt in March

1985, and fertilized (N, P or N+P) eight months later. In situ measurements

of N mineralization and nutrient mobility were compared with those from an

adjacent, undisturbed forest. Nitrification was absent in all soils.

Nitrate concentrations in soil water were low and probably not responsible

for cation movement. Cation concentrations in soil water were immediately

increased by fire and by fertilizer additions and decreased continuously

after these events. Addition of strong-acid anions in fertilizer increased

cation movement. Soil water sampled at 10 cm depth was often coloured and

contained high concentrations of oxidizable organic carbon; lower

concentrations were found in samples collected from 30 cm depth. Uptake and

mineralization of N were increased by logging and slash-burning.

N-mineralization was promoted further by the addition of fertilizer-N and

was reduced by the addition of P, or N+P. Nutrient losses were exacerbated

by fertilizer additions and restricted by lack of nitrification, by

nutrient uptake by soil microorganisms and vegetation, and probably by

formation of complexes between organic anions and cations in the soil

profile.

 

Adams, M. and P. Attiwill (1991). "Nutrient balance in forests of northern

Tasmania. 1. Atmospheric inputs and within-stand cycles." Forest Ecology

and Management 44(2-4): 93-113.

Six undisturbed forests were studied: dry or wet sclerophyll (dominated by

Eucalyptus obliqua, E. amygdalina, E. regnans) and cool temperate rain

forest (principal species Atherosperma moschatum, Nothofagus cunninghamii).

Forest soils were ammonifying and had low concentrations of extractable P,

inorganic N and cations, irrespective of geology. Litterfall was

proportional to productivity, could be described by a climatic index, and

nutrient return via litterfall was the dominant transfer for N, P and Ca.

However, significant amounts of N (up to 8 kg/ha per year) and other

nutrients were added via rainfall, and total nutrient input was related to

rainwater quality. Soil in the rain forest was more organic than in the

eucalypt forests, and had greater amounts of P, N etc. Litterfall in the

rain forest was rich in P, but poor in Mg, relative to the eucalypt

forests, and nutrient absorption by the root mat was more effective than

that in the eucalypt forests.

 

Attiwill, P. M. (1986). "Interactions between carbon and nutrients in the

forest ecosystem." COUPLING OF CARBON, WATER AND NUTRIENT INTERACTIONS IN

WOODY PLANT SOIL SYSTEMS. PROCEEDINGS OF A SYMPOSIUM OF THE INTERNATIONAL

UNION OF FORESTRY RESEARCH ORGANIZATIONS. Luxmoore, R.J. 2: 1-3.

The sustained supply of nutrients in forest ecosystems depends on

processes by which nutrients are cycled from plant (in organic

combinations) to soil and back to plant (in simple inorganic form). Studies

of the key processes of decomposition and mineralization, and of equilibria

determining nutrient availability have been hampered by lack both of

appropriate chemical methods and of methods that distinguish among

fractions of organic matter of varying nutritional quality. The root

systems of forests must also be studied more intensively. In particular,

mechanisms by which nutrients in short supply are taken up (for example,

the role of mycorrhizae and of specialized systems such as proteoid roots)

and the redistribution of nutrients associated with turnover of the fine

root system are fields for future research.

 

Attiwill, P. M. and M. A. Adams (1993). "Tansley Review No. 50. Nutrient

cycling in forests." New Phytol 124(4): 561-582.

Studies of nutrient cycling in forests span more than 100 yr. In earlier

years, most attention was given to the measurement of the pools of

nutrients in plants and soil and of the return of nutrients from plant to

soil in litterfall. The past 20 yr or so have seen a major concentration on

the processes of nutrient cycling, with particular emphasis on those

processes by which the supply of nutrients to the growing forest is

sustained. In the more highly productive forests, up to 10 tonnes of litter

of low nutritional quality is deposited annually on the forest floor. The

decomposition of this litter, the mineralization of the nutrients it holds,

and the uptake of nutrients by tree roots in the carbon-rich environment

which results are the themes of this review. Studies of decomposition of

litter in forests have been dominated by the role of nitrogen as a limiting

factor, a domination which reflects the preponderance of studies of

temperate forests in the Northern Hemisphere. For many forests of the world

growing on soils of considerable age, it seems more probable that growth

and nutrient cycling are limited by phosphorus (or some other element).

There is increasing evidence for a number of forests that phosphorus is

immobilized in the first stages of decomposition to a significantly greater

extent than is nitrogen. Advances in research will depend, as with studies

of soil organic matter, in defining and developing analytical techniques

for studying biologically active forms of potentially limiting nutrients,

rather than total elemental concentrations. Net nitrogen mineralization

rates approximate rates of nitrogen return in litterfall but the

contribution of nitrification is variable. Nitrification is not inhibited

by the low pH of many forest soils and there is increasing evidence of

nitrate immobilization by microorganisms and of increased diversity and

better competitiveness for NH sub(4) super(+) of nitrifying microorganisms

than has previously been accepted. Variability in rates of nitrification is

often interpreted as being due to allelopathy. Hypotheses invoking

allelopathy are more or less untestable, and it seems likely that new

techniques using super(15)N in situ will lead to a more fundamental

understanding of nitrogen transformations in forest soils. Recent studies

in coniferous forest soils have highlighted the short (< 1 d) turnover time

of NH sub(4) super(+).

 

Bowman, D. M. J. S., A. R. Maclean, et al. (1986). "Vegetation-soil

relations in the lowlands of south-west Tasmania." Aust. J. Ecol. 11(2):

141-153.

A geographic survey of 14 south-west Tasmanian sedgeland-heaths revealed

that soil organic matter is related to: water content; total nitrogen (N);

total and exchangeable sodium (Na), calcium (Ca) and magnesium (Mg);

exchangeable potassium (K) cation exchange capacity: and total exchangeable

bases. However, total and available phosphorus (P), total K and iron (Fe),

pH level and percentage base saturation were found to be independent of

organic content. Most of the soil nutrient capital is contained in the A

sub(0) horizon, the depth of which was found to be positively related to

the time elapsed since the last fire. There is no clear relationship

between rock type and soil fertility, but there is evidence of

soil-vegetation interaction. The sedgeland-heath species have lower

concentrations of P, Ca and Mg in their foliage and are more efficient in

the withdrawal of P and K upon tissue senescence than the surrounding scrub

and forest species. Over a vegetation transition from sedgeland-heath to

forest on uniform geology there was a change in soil type.

 

Campbell, E. O. (1983). "Mires of Australasia." MIRES: SWAMP, BOG, FEN AND

MOOR. REGIONAL STUDIES. Gore, A.J.P. ed.

The author reviews the wetlands of Australia, Tasmania and New Zealand;

and the effects of climate on their development and geographical

distribution. Wetland vegetation are covered and used to classify types of

wetlands as well as soil composition. The main land masses of the

Australasian region are Australia, Tasmania and New Zealand. Australia is

an island continent with an area over three-quarters that of Europe.

Politically it is divided into the Australian Capital Territory, the

Northern Territory and the states of Western Australia, Queensland, New

South Wales, Victoria and South Australia. Tasmania is an island lying some

200 km to the south of Victoria. It is politically part of Australia and

has an area slightly less than that of Scotland. New Zealand lies some 1600

km to the southeast of Australia. It consists of three main islands, known

as North Island, South Island and the smaller Stewart Island, with a total

area about seven-sixths of that of Great Britain.

 

Chambers, D. and P. Attiwill (1994). "The ash-bed effect in Eucalyptus

regnans forest: chemical, physical and microbiological changes in soil

after heating or partial sterilisation." Australian Journal of Botany

42(6): 739-749.

The ash-bed effect (the enhanced growth of plants on soil which has been

heated) following fire in Eucalyptus regnans forest is dramatic. The

results are presented of studies of the effects of a range of heating and

partial sterilization treatments on chemical, microbiological and physical

properties in soil from a 250-year-old E. regnans forest in Victoria. Soil

treatments not involving heat (chemical sterilization, gamma-irradiation

and air-drying) and the lower temperature heat treatments (100 and 200ƒC)

had no marked effects on physical characteristics. All treatments produced

more or less similar effects on microbial populations. On the other hand,

heating the soil to 400-600ƒC produced large, significant and sustained

increases in the availability of nitrogen and phosphorus and these

increases were enhanced by a decrease in clay colloid. The results support

the hypothesis that the ash-bed effect following fire in E. regnans forest

is due to an increase in the availability of nutrients, and in the

availability of nitrogen and phosphorus in particular. A transitory

increase in the concentration of manganese caused by heating the soil may

account for initial toxicity in plants grown in soils which have been

heated. Since species within the subgenus Monocalyptus are characterized by

lower tissue concentrations of manganese than those within Symphyomyrtus,

it could be hypothesized that the potential for toxicity following bushfire

varies between the two subgenera. The literature on the effects of

soil-sterilising treatments is highly variable; the causes of variability

include soil type and moisture content, treatment (sterilizing by steam,

chemicals or heat) and the method of treatment (time, how the soil was

contained, and how the treatment was applied).

 

Colhoun, E. a. (1992). "Late glacial and Holocene vegetation history at

Poets Hill Lake, western Tasmania." Australia Geographer 23(1): 11-23.

Two cores (down to 3.9 m) were taken from the edge of a peaty flat by

Poets Hill Lake (600 m altitude), Tasmania, for pollen analysis to give a

record of vegetation history since the last glaciation (Margaret

Glaciation). Alpine herbfield, coniferous heath and Nothofagus gunnii scrub

developed on the moraine until 11 400 BP. Wet montane forest and heath then

developed with Phyllocladus aspleniifolius, N. cunninghamii and Eucalyptus

until about 10 000 BP. After 10 000 BP, there was a mosaic of N.

cunninghamii rain forest, Myrtaceae and Proteaceae scrub and Sprengelia

incarnata heath. The development of the vegetation from alpine communities

to temperate rain forest, which is near its limit at 600 m altitude,

occurred under the influence of improving climatic conditions with rapid

upslope migration or local expansion of taxa during the late glacial.

Temperatures were warm enough for the development of rain forest at 600 m

altitude by 10 000 BP, if not earlier. The development of a mosaic of rain

forest, scrub and heath vegetation rather than extensive rain forest after

10 000 BP reflects the influence of poor soils, poor drainage and fire.

Comparison with similar pollen diagrams from W. Tasmania suggests that the

development of pollen/vegetation associations was time transgressive with

altitude during the late glacial when climatic influences and migration

rates were important, and that the mosaic of vegetation communities became

more complex during the Holocene because of adjustment to or control by

local ecological factors.

 

Davidson, N. J. and J. B. Reid (1989). "Response of eucalypt species to

drought." Aust. J. Ecol no. 2: pp.

The response of three eucalypt species (Eucalyptus pulchella, Eucalyptus

coccifera and Eucalyptus delegatensis ) to a severe drought in the summer

of 1982/83 was examined at Snug Plains, south-eastern Tasmania. Few large

differences in leaf water potential ( psi sub(l)) or stomatal conductance

(gs) were apparent even at the height of the drought when both psi sub(l)

and soil water potentials ( psi sub(s)) reached ca. -4.5MPa.

 

Duncan, F. and Kiernan K (1989). "Drought damage in a Tasmanian forest on

limestone." Helictite 27(2): 83-86.

Widespread but patchily distributed drought death of forest trees (both

overstorey and understorey species) occurred in early 1988 on a limestone

ridge at Mole Creek, Tasmania. Moderate to severe symptoms of drought

stress were observed in Eucalyptus viminalis, E. delegatensis, E. obliqua,

Pomaderris apetala, Olearia argophylla, O. lirata, Cassinia aculeata,

Pultenaea juniperina and Senecio linearifolius while Bursaria spinosa

showed no drought symptoms and Acacia melanoxylon was only partly affected.

The close occurrence of damaged and undamaged vegetation (e.g.

juxtaposition of severely damaged specimens of the xerophytic species

Pultenaea juniperina with healthy specimens of the wet sclerophyllous

species Pomaderris apetala) is thought to reflect differences in the speed

of soil moisture decline down the length of individual soil-filled solution

tubes in which the trees were rooted. Possible palaeoecological,

geomorphological and silvicultural implications are briefly discussed.

 

Ellis, R. C. and A. M. Graley (1987). "Soil chemical properties as related

to forest succession in a highland area in north-east Tasmania." Aust. J.

Ecol. 12(3): 307-317.

The relationship between vegetational type and a number of soil chemical

factors was examined in secondary successions from fire-maintained

eucalypt/grass to climax rainforest communities growing on uniform granitic

soil parent material. Canonical variates analysis, which utilized the

following variables: pH; loss on ignition; total N, P, Ca, and Mg; cation

exchange capacity and exchangeable Ca, K, and Mg; and potentially

mineralizable N, revealed close overall similarity between surface soils of

adjacent types, and significant differences among those of types distant

from each other in the successional sequences. Differences among soils in

chemical composition and rates of mineralization of N were due to

differences in species composition of the vegetational types that they

carried for the time being.

 

Ellis, R. C. and P. I. Pennington (1989). "Nitrification in soils of

secondary vegetational successions from Eucalyptus forest and grassland to

cool temperate rainforest in Tasmania." Plant Soil 115(1): 59-73.

Rates of nitrification in well drained granitic soils from forest stands

and grassland of differing successional status and from beneath isolated

individuals of several tree species were compared in a series of laboratory

experiments. Nitrification was absent or occurred at only a low rate in

many soils; it generally increased as succession proceeded from mature

grassland or eucalypt forest towards climax temperate rainforest, but

decreased in mature climax forests. However, the influence of individual

tree species was often paramount. Nitrification was stimulated by

disturbance of a stand by disease. A possible inhibitor of nitrification in

a rainforest soil could not be removed by leaching with water, nor

transferred via the leachate to a nitrifying soil. Addition of P was

without effect on either total amount of nitrate produced or on net

mineralisation of soil N, but sometimes increased the rate of nitrification

of added ammonium.

 

Ellis, R. C. and P. I. Pennington (1992). "Factors affecting the growth of

Eucalyptus delegatensis seedlings in inhibitory forest and grassland

soils." Plant Soil no. 1: pp.

In many highland forests of Eucalyptus delegatensis in Tasmania the

establishment and healthy growth of eucalypts is promoted and maintained by

fire. In the absence of fire, secondary succession from eucalypt forest to

rainforest occurs, during which the eucalypts decline and die prematurely.

On sites that are prone to radiation frost severe reduction or removal of a

tree canopy allows a sward of tussock grasses to develop, in competition

with which seedlings of eucalypts decline in growth and a high proportion

dies. Factors of the soil that could contribute to these phenomena were

investigated by means of pot experiments that used soils from: a secondary

succession of vegetative types from recently burned healthy eucalypt forest

to unburned mature rainforest: this encompassed a sequence of decline and

death of the eucalypt trees; soil from old grassland in which eucalypt

seedlings were exhibiting severe growth check and mortality; from beneath

individual trees of several species growing on old grassland.

 

Fensham, R. J. (1989). "The pre-European vegetation of the Midlands,

Tasmania: A floristic and historical analysis of vegetation patterns." J.

Biogeogr. 16(1): 29-45.

83% of the area of native vegetation of the Midlands has been replaced

after 170 years of agricultural exploitation. However, scattered remnants

of the native vegetation still exist today in most Midlands environments,

and these were surveyed. The floristic variation within and between these

remnants was most closely related to soil moisture, as affected by texture

and topography. A classification produced thirteen groups that comprised

subdivisions of grassy communities dominated by Eucalyptus amygdalina

Labill. forest, E. viminalis Lag., E. pauciflora Sieber ex Spreng., E.

ovata Labill. woodlands, and treeless communities.

 

Fensham, R. and J. Kirkpatrick (1992). "The eucalypt

forest-grassland/grassy woodland boundary in central Tasmania." Australian

Journal of Botany 40(2): 123-138.

Downslope boundaries of forest with grassland and grassy woodland occur

over a wide altitudinal range in central Tasmania. Observations were made

and a series of experiments were carried out at 3 sites to study the causes

of these boundaries at low, medium and high altitudes (500, 800 and 1000 m,

respectively). Open vegetation was generally associated with moister and

less rocky soils and more subdued topography than the adjacent forest.

Frost incidence and intensity, soil moisture and waterlogging varied

markedly among the 3 open areas. Direct sowing trials were attempted at

different times using various ground treatments and species mixtures of

woodland eucalypts (Eucalyptus ovata, E. rodwayi and E. gunnii) and forest

eucalypts (E. tenuiramis, E. pauciflora and E. coccifera ). Seedlings

survived 4 years in the open at all sites, and seedlings established in the

open both naturally, and after sowing, where grass competition was reduced

by herbicide application, scarification or root competition from adult

eucalypts. Grazing had no detectable effect on seedling establishment. A

pot experiment demonstrated a suppressive effect of native grass (Poa

labillardieri) swards on establishment and growth of E. rodwayi seedlings;

this effect was largely independent of available moisture and nutrients.

While frost, waterlogging, fire and drought may play a role in inhibiting

eucalypt establishment and increasing eucalypt mortality at some or all of

the sites, the dense grass swards found in all the open areas are

considered to be the most likely primary agent of tree exclusion.

 

Frey, W. and P. Dalton (1996). "Hypnodendron comosum-community in

Tasmania." Nova Hedwigia 62(1-2): 215-220.

Hypnodendron comosum var. comosum var. sieberi and H. vitiense subsp.

australe (endemic in SE Australia, Tasmania and New Zealand and thought to

be archaic taxa) form a typical dendroid community. The dendroid taxa reach

high cover values and characterize the feature of the community ("miniature

forest"), which is described for the first time from Tasmania. Surveys were

made in western Tasmania, in rain forests characterized by Nothofagus

cunninghamii, Atherosperma moschatum and Eucryphia lucida and the conifers

Phyllocladus, Lagarostrobus [Lagarostrobos] and Athrotaxis. On loamy and

humic soil the number of species of the community is low, increasing on

decaying wood. The community is a complementary one to those described from

New Zealand. It is assumed that such dendroid bryophyte communities from

forests which today are temperate rain forests of the southern hemisphere

represent a relict vegetation of the forest floor of former Gondwanaland

forests.

 

Gibson, N. and J. B. Kirkpatrick (1985). "Vegetation and flora associated

with localized snow accumulation at Mount Field West, Tasmania." Aust. J.

Ecol. 10(2): 91-99.

The vegetation associated with a snow patch at Mt Field in Tasmania is

described and mapped. Seven distinct vegetation types were found to be

related directly to topography and hence to the patterns of snow

accumulation, snow melt and soil drainage. The fjaeldmark found where snow

lies longest in unusual for Tasmania and may be the product of past

climatic events and a peculiar parent material. At the generic level there

exists a high similarity with the snow patch flora of the Australian Alps.

 

Gibson, N. (1990). "The environments and primary production of cushion

species at Mt Field and Mt Wellington, Tasmania." Aust. J. Bot no. 3: pp.

Primary production of four species of alpine cushion plants were studied

over a 2-year period. The climate of these areas was found to be severe but

with a high degree of variability on a seasonal and yearly basis. The

growing season at the higher altitude sites generally exceeded 6 months.

Net above ground primary production of the four cushion species ranged from

282 to 709 g m super(-2) year super(-1). Reproductive effort fluctuated

between species and years, ranging from 0 to 30% of net above ground

production. Patterns in dry matter accumulation suggest no individual

species would show consistently superior growth rates under present

climatic conditions. Soil moisture and soil nutrient status was found to be

similar between all sites. Altitude of the sites (830-1400 m) was found to

be strongly correlated with the timing of flowering and/or seed set but

appeared to have little effect on net primary production.

 

Gibson, N., Brown Mj, et al. (1992). "Flora and vegetation of ultramafic

areas in Tasmania." Australian Journal of Ecology 17(3): 297-303.

The flora and plant communities were surveyed of 15 areas of ultramafic

(serpentine) rock in Tasmania. Dry sclerophyll communities on ultramafic

rock were restricted to 2 areas in the north of Tasmania. In the wetter

western and southern areas the ultramafic land supported heathlands, wet

scrubs, and a wet sclerophyll/mixed forest community dominated by

Eucalyptus obliqua, E. nitida or E. delegatensis. One of the most striking

features of the ultramafic areas in Tasmania is the variable degree of

physiognomic differentiation between adjacent ultramafic and non-ultramafic

substrates at the different sites. Physiognomic differentiation is less on

non-lateritic soil types or where accumulation of colluvial material has

occurred. Only 3 ultramafic endemic taxa were recorded.

 

Grant, J., Laffan M, et al. (1995). "Mapping State Forest soilsin Tasmania." ACLEP Newsletter 4(2): 16-19.

Methods of soil mapping and characterization for the Tasmanian State

Forest at scales of between 1:50 000 and 1:100 000 are outlined. The

results are entered in a database and digitized soil boundaries in a

geographical information system. This information can be used to produce

soil/vegetation/forest productivity models for projection onto the

landscape.

 

Grant, J., Laffan Md, et al. (1995). "Forest soils of Tasmania: a handbook

for identification and management." 189(5).

This book describes a range of forest soils commonly found in Tasmania in

terms of their site and profile characteristics. It provides interpretation

of potential degradation hazards for each soil and recommendations for

management. Colour photographs of typical soil profiles together with

associated native vegetation are included as an aid for identification. A

glossary of soil terms is included, as well as many annexes giving detailed

information and analysis of the soils, and listing the plant species

mentioned in the site descriptions.

 

Grierson, P. and P. Attiwill (1989). "Chemical characteristics of the

proteoid root mat of Banksia integrifolia L." Australian Journal of Botany

37(2): 137-143.

The proteoid roots of B. integrifolia are concentrated in the topsoil,

forming a dense mat beneath the litter layer. Water extracts of the

proteoid root mat of a mature B. integrifolia var. cunninghamii stand in

Victoria contained a significantly greater amount of hydrogen ions,

reducing agents and an unidentified chelating agent than water extracts of

soil beneath the root mat, of the litter layer or of soil from beyond the

proteoid root zone. The results are discussed in relation to previously

reported production of organic chelates by plants causing solubilization of

soil phosphates. It is suggested that proteoid roots of B. integrifolia

chemically modify the soil environment, thereby enhancing nutrient uptake.

 

Horne, R. and J. Hickey (1991). "Review. Ecological sensitivity of

Australian rainforests to selective logging." Australian Journal of Ecology

16(1): 119-129.

A review of past studies of the ecological consequences of selective

logging in the major rain forest areas of Australia. The main aspects

covered are recovery of stand structure, regeneration capacity, individual

tree growth and species composition of stands, hydrological effects,

floristics, wildlife, soil nutrient levels, fire susceptibility, and

incursion of weeds and diseases. Following a single selective logging, the

changes indicated by individual studies often appeared to be relatively

minor. It is suggested that many of these effects are not extensive or

irreversible and might not persist beyond structural recovery of the rain

forest. However, two changes were identified as likely to persist beyond

structural recovery. These are (i) a post-logging difference in the

proportional representation of major overstorey tree species and (ii) a

reduction in the numbers of large-diameter trees. More extensive and

longer-lasting changes may result from multiple selective loggings,

especially if the interval between loggings is short. Even for light

logging intensities, a conservative interval of at least 60 years between

loggings is indicated, to allow canopy and below-canopy conditions to be

restored. The slower growth of the rain forests in Tasmania, compounded by

a geographical susceptibility to drought, increases the possibility of fire

damage following selective logging relative to the more northerly mainland

rain forests.

 

Judd, T. S., L. T. Bennett, et al. (1996). "The response of growth and

foliar nutrients to fertilizers in young Eucalyptus globulus (Labill.)

plantations in Gippsland, southeastern Australia." For. Ecol. Manage. 82:

1-3.

Fertilizer trials, which included rates of application of up to 400 kg/ha

N, 200 kg/ha P and 200 kg/ha K, were established in experimental

plantations of E. globulus at three sites in Gippsland, southeastern

Australia. These sites cover a range of annual rainfall and soil fertility,

from 620 mm on a deep infertile sand, to 1000 mm on a rich gradational clay

loam. Early growth at all sites was significantly increased by the addition

of fertilizers and by age 4 years was consistently best at the highest

combined rates of N and P. Foliar N and P concentrations were significantly

increased at all sites by combined additions of N and P at Age 1, but were

insensitive to treatment at Ages 2 and 4. Relationships between foliar

nutrient concentrations and growth were site dependent. Correlations

between foliar nutrients and growth in the same year were strongest at Age

1. Foliar nutrients at Age 1 were also strongly associated with growth at

later ages at all sites. The best growth was associated with foliar

concentrations at Age 1 of about 2.5% N and 0.23% P. Additions of P, alone

or with N, consistently decreased foliar N/P ratios at Age 1 to between 11

and 12 whereas N and P additions resulted in foliar N/P ratios of 15 to 16

at Age 4. These ratios indicate a greater requirement for P than N in the

first year suggesting that the N/P ratio of fertilizers should be 1:1

within the first year, increasing to 2:1 in later applications. General

recommendations for fertilizer addition in the routine establishment of E.

globulus are presented. Mean tree volumes at Age 4 ranged from 0.014 to

0.019 m3 in control treatments and from 0.031 to 0.055 m3 at the highest

rate of fertilizer addition. Growth responses to fertilizers were greatest,

in both absolute and relative terms, at the most fertile site and appeared

to be constrained by unfavourable soil texture and limited water

availability at the other sites. Because the responses of E. globulus are

site specific, detailed assessments of fertilizer requirements on soil

types other than those described here require the establishment of further

trials. A schedule for the establishment and monitoring of such trials is

proposed.

 

Kirkpatrick, J. B. (1983). "Treeless plant communities of the Tasmanian

High Country." MOUNTAIN ECOLOGY IN THE AUSTRALIAN REGION. Purdie, R.W. 12:

61-77.

Analyses of the associations of 65 dominant species from 430 quadrats

located in Tasmanian treeless high altitude vegetation are used to provide

a framework and guidelines for the construction of a typology of plant

communities. These communities are listed and discussed within the context

of the following vegetation types: bolster heath, deciduous heath,

coniferous heath, heath, fjaeldmark, bog, fen, short alpine herbfield, tall

alpine herfield and tussock grassland. The distribution of communities is

best related to a climatically and geologically-controlled edaphic

gradient, a soil drainage gradient and to the vagaries of fire history. The

successional status of most of the plant communities is deduced from their

patterns of distribution. Several of the alpine dominants usually fail to

regenerate after fire. There is insufficient evidence to support a widely

suggested cyclic succession process involving bolster plants.

 

Kirkpatrick, J. B. and K. J. M. Dickinson (1984). "The impact of fire on

Tasmanian alpine vegetation and soils." Aust. J. Bot no. 6: pp.

Observations were made across 11-40-year-old fire boundaries in Tasmanian

alpine areas of varying macroenvironment and flora. Surface soil organic

matter and total nitrogen were significantly less where the vegetation had

been recently burned. There were no significant differences between

recently burned and recently unburned plots for phosphorus, potassium,

calcium, sodium and pH. The burned plots contained few or no gymnosperms or

deciduous shrubs, the most frequent dominants of the unburned vegetation.

Most other shrubs were markedly less important in the burned than in the

unburned plots, although most species of bolster form were little affected

by fire, and some composite shrubs were most abundant on the burned plots.

Most herbaceous species had equal or higher cover on the burned plots than

on the unburned plots. The burned vegetation of the eastern mountains

appeared to regenerate more quickly than that of the more oligotrophic

western mountains.

 

Kirkpatrick, J. B. (1986). "Conservation of plant species, alliances and

associations of the treeless high country of Tasmania, Australia." Biol.

Conserv. 37(1): 43-58.

The treeless high country of Tasmania has considerable conservation and

catchment value. Much of its area lies within National Parks and other

reserves of similar tenure, but many species and communities are confined

to land of less secure status. Much high mountain land is subject to

stockgrazing and burning, both of which activities have led to

deterioration in vegetation and soils. The reservation status of species

and communities can be made near perfect with relatively small additions to

the State Reserve system. However, the impacts of trampling and deliberate

fires do not respect reserve boundaries, and their mitigation or

elimination requires some intensive management.

 

Kirkpatrick, J. B. and F. Duncan (1987). "Tasmanian high altitude grassy

vegetation: Its distribution, community composition and conservation

status." Aust. J. Ecol. 12(1): 73-86.

Grassy woodland, grassy shrubland, grassy sedgeland, tussock grassland and

grassland are extensive on basalt, limestone and fine-textured Quaternary

deposits, are occasional on dolerite, granite and fine-grained sedimentary

rocks, but are absent from the siliceous mountains of Tasmania. With the

exception of limestone lithosols, the grassy communities are confined to

relatively deep soils with a low surface rock cover. Much of the area of

the grassy communities below the climatic treeline has clearly been forest

in the recent past, although some of the higher subalpine plains seem

likely to have been grassy at least since the peak of the Last Glacial.

 

Ladd, P., D. Orchiston, et al. (1992). "Holocene vegetation history of

Flinders Island." New Phytologist 122(4): 757-767.

Two swamp sites on Flinders Island in Bass Strait, Tasmania, provide

evidence of vegetation cover for the period 10 000 BP to present. Steppe

vegetation in which Compositae Liguliflorae taxa and chenopods were

important was present on the Flinders Island part of the Bassian Isthmus

during the earliest part of the record. However, it was replaced by

eucalypt forest or woodland with a grassy understorey and some shrubs as

sea level rose to form the present island by 6000 BP. Eucalypt dominated

vegetation became less important about 940 BP when Callitris became

prominent until very recently. This change may be related to a drier

climate. From pollen evidence in this study and that from other SE mainland

and Tasmanian sites it is suggested that apparent prominence of Casuarina

in SW Victoria and SE South Australia during the early Holocene was due to

local soil factors and drier climate. Later changes in soil and climate led

to a decrease in Casuarina and increase in Eucalyptus.

 

Podger, F. and M. Brown (1989). "Vegetation damage caused by Phytophthora

cinnamomi on disturbed sites in temperate rainforest in western Tasmania."

Australian Journal of Botany 37(6): 443-480.

P. cinnamomi was isolated directly from 558 diseased plants of 39 species,

including 1 fern, 4 graminoids and 34 woody dicotyledons, all indigenous to

the cool temperate rain forest of Tasmania. Pathogenicity was proved by

greenhouse tests (20 spp.) and by field inoculation at two sites (19 spp.).

Of the 142 species in the rain forest flora, 69 were rated for the

susceptibility of field populations: 30% were highly susceptible and less

than 5% highly resistant. High proportions of susceptible species occurred

in Epacridaceae, Eucryphiaceae and Proteaceae. All 93 isolates tested were

the common A2 mating type. The fungus was consistently associated with

disease at 47 survey sites. Diseased plants were widely but patchily

distributed along exposed road and track edges within unburned rain forest

and in recently burned rain forest. The fungus was not recovered from soil

samples taken beneath healthy roadside regeneration, beneath undisturbed

rain forest or above 900 m alt. The disease has the characteristics of

attack by a recent invader and appears to be dependent upon disturbance

which increases soil temp. above 15ƒC, the lower threshold for infection by

P. cinnamomi. Post-fire recovery of forest canopy is expected to allow

re-establishment from external sources of seed of those susceptible species

which are both efficiently dispersed and tolerant of shade. Species lacking

such characteristics are at risk of local elimination but no single rain

forest species appears to be under threat of extinction from this pathogen.

 

Polglase, P. J., P. M. Attiwill, et al. (1986). "Immobilization of soil

nitrogen following wildfire in two eucalypt forests of south-eastern

Australia." Acta Oecol. Oecol. Plant. 7(3): 261-271.

Forests of Eucalyptus obliqua (Messmate) and E. regnans (Mountain Ash)

were burnt by crownfire in November of 1982. Litterfall increased 2-fold in

the 3 months following the fire, about 63% of which was scorched dead

leaves. Concentrations of N and P in these scorched leaves were about twice

those in leaf litter of unburnt forests. Thus in the first year after the

fire, total N and P in litterfall was 2.5 times greater than normal in

Mountain Ash and 1.5 times greater than normal in Messmate. Concentrations

of N and P in leaf litter of Messmate (which regrew from epicormic buds)

returned to normal after 12 months. Available P and total inorganic N in

the soil increased significantly after the fire. Inorganic N in the soil

beneath Messmate is normally NH sub(4)@)u+-N; even after intense fire, no

NO sub(3)@)u--N was produced. The increase in inorganic N was rapidly

immmobilized and returned to normal levels within 5 months.

 

Polglase, P. and P. Attiwill (1992). "Nitrogen and phosphorus cycling in

relation to stand age of Eucalyptus regnans F. Muell. I. Return from plant

to soil in litterfall." Plant and Soil 142(2): 157-166.

The N and P contents of the litter layer and the return of these nutrients

in litterfall were measured in seven stands of mountain ash (Eucalyptus

regnans) in Victoria, Australia, ranging in age from 5 years to _ 250

years. Both annual litterfall and nutrient return were correlated with

stand basal area and were high compared with other productive eucalypt

forests. In contrast, the fall of dead eucalypt leaves was constant with

stand age, demonstrating that sites are fully occupied at an early age.

Similarly, amounts of N and P in total leaf fall (overstorey plus

understorey) were constant with stand age, except for low amounts in the

stand aged 40 years where Acacia spp., important fixers of atmospheric N,

were not prevalent. The decomposition constant (k) of organic matter in the

litter layer decreased with stand age, from 0.3/year at age 5 years to

0.23/year at age 250 years. These constants also applied to N and P,

indicating a tight coupling between organic matter decomposition and

release of these nutrients from litter. The litter layer released _ 30

kg/ha of N at age 5 years, and _ 70 kg/ha at age 80 years. These results

are discussed in relation to growth of mountain ash following fire, and the

subsequent retention and accumulation of N during stand development.

 

Polglase, P., P. Attiwill, et al. (1992). "Nitrogen and phosphorus cycling

in relation to stand age of Eucalyptus regnans F. Muell. II. N

mineralization and nitrification." Plant and Soil 142(2): 167-176.

Laboratory and in situ rates of N mineralization in soils from mountain

ash (Eucalyptus regnans) forests in Victoria, Australia, were measured

fortnightly for two years in stands aged 9, 40, 80 and _ 250 years

(overmature), and for one year in stands aged 5 and 46 years. Rates of

anaerobic mineralization (the laboratory test) showed little seasonal or

annual variation. In contrast, rates of in situ mineralization varied

markedly with season, being highest in spring and summer. Anaerobic

mineralization was highly correlated with stand age to 80 years, but

decreased between ages 80 and 250 years. In situ mineralization also

decreased between these two ages, but otherwise was not related to stand

age. Hence, the correlation between anaerobic and in situ mineralization

along the age sequence was weak, suggesting that the anaerobic test is of

maximum utility when this pool is in balance with inputs from decomposing

litter ('steady-state' ecosystems). Nitrification was strong in stands aged

9, 46, 80 and 250 years and weak in stands aged 5 and 40 years. Within

stands, the rate of nitrification during each period of in situ containment

was highly dependent on the supply of NH4 ions. Between stands, annual

rates of nitrification appear to be related to the balance between the N

capital of the site, its rate of turnover, and the demand for N by

heterotrophs and vegetation so that if NH4 supply is depleted, little is

left for autotrophic nitrifiers.

 

Polglase, P., P. Attiwill, et al. (1992). "Nitrogen and phosphorus cycling

in relation to stand age of Eucalyptus regnans F. Muell. III. Labile

inorganic and organic P, phosphatase activity and P availability." Plant

and Soil 142(2): 177-185.

Concentrations of labile inorganic P (Pi, dilute double acid-extractable),

labile organic P (Po, bicarbonate-extractable), microbial P, and

phosphatase activities were measured in mountain ash (Eucalyptus regnans)

stands aged 0, 5, 9, 16, 40, 46, 80 and 250 years in Victoria, Australia.

Labile Pi, initially 34 µg/g at age 0 years, decreased to 2.3 µg/g at age

16 years, and was constant thereafter. All other indices of available P

increased with stand age. Microbial P varied the most along the age

sequence, from 1.8 µg/g at 0 years to 99 µg/g at age 80 years. Oxidation of

organic P during the regeneration burn creates a large and readily

available source of Pi for the young, developing stand. After canopy

closure, the stand becomes increasingly dependent on P released by

mineralization. Phosphatase activities were as high as have been recorded,

and the organic pools are therefore significant stores of potentially

available P. At age 80 years (for example) the amount of labile Po and

microbial P in the surface 5 cm of soil was 26 kg/ha. However, the

adsorption maxima for Pi fixation were also as high as has been recorded,

and so it is apparent that roots must possess specialized mechanisms of

uptake to compete for mineralized P.

 

Potts, B. M. and J. B. Reid (1990). "The evolutionary significance of

hybridization in Eucalyptus." Evolution 44: 2151-2152.

New estimates of dispersal and levels of hybridization were obtained from

open-pollinated progenies from extra sites and from post-dispersal,

pre-selection seed samples collected from ants' nests and soil from the

forest floor immediately after fire to support the authors' earlier

conclusions that for E. risdonii pollen-mediated dispersal into the range

of E. amygdalina occurs over a greater distance than seed-mediated

dispersal. Recent criticisms of the subject [Schemske, D.W.; Morgan, M.T.

(1990) Evolution 44, 2150-2151] are answered.

 

Read, J. and R. S. Hill (1983). "Rainforest invasion onto Tasmanian

old-fields." Aust. J. Ecol no. 2: pp.

The regeneration of rainforest onto land cleared for grazing early this

century was studied on several sites in northern Tasmania. Drimys

lanceolata , a bird-dispersed species, was the main invader. The climax

forest species, Nothofagus cunninghamii and Atherosperma moschatum were

invading slowly from the forest edge with occasional trees established in

the field. Woody plants in the old-field were clumped around logs. This was

related to the role of logs in attracting seed and to possible roles as

competition-free sites and sites safe from browsing and climatic stresses.

Changes in dominance by particular life forms appeared to be related to

dispersal events, environmental modification by the developing vegetation

and life history characteristics. The extremely slow invasion by climax

species is due to the absence of bare mineral soil as well as to dispersla

characteristics, browsing and possibly exposure to climatic stresses.

 

Read, J. (1995). "The importance of comparative growth rates in determining

the canopy composition of Tasmanian rainforest." Australian Journal of

Botany 43(3): 243-271.

Growth rates of the rain forest canopy species Atherosperma moschatum,

Eucryphia, Nothofagus cunninghamii, Athrotaxis selaginoides and

Phyllocladus aspleniifolius were measured in naturally-occurring seedlings

and in seedlings grown in greenhouse pot trials. The highest field growth

rates were recorded in N. cunninghamii on lowland sites in fertile,

well-drained soils. On poorer soils, growth rates of N. cunninghamii were

lower and there was little difference in growth rates between species. On

the most acidic and probably infertile soil, the growth rates of P.

aspleniifolius was 1.4 times greater than that of N. cunninghamii, although

not significantly different at the 95% level. Similar trends in growth

rates were recorded in pot trials. High growth rates were recorded in N.

cunninghamii and E. lucida on fertile, well-drained soils, with generally

lower growth rates in Atherosperma moschatum, P. aspleniifolius and

Athrotaxis selaginoides. No significant differences in growth rates was

recorded among N. cunninghamii, E. lucida, Atherosperma moschatum and P.

aspleniifolius when grown in poor soils or in low nutrient treatments. The

trends in comparative growth rates correlate with the measured canopy

composition of the study sites. Nothofagus cunninghamii dominates forest

stands on sites where it has a clearly superior growth rate compared with

co-occurring light-demanding species. Mixed canopies occur on sites where

there is little or no difference between species' growth rates. However,

not all patterns of canopy dominance can be explained by comparative growth

rates alone. For example, P. aspleniifolius commonly dominates forest

stands at low- to mid-altitude where growth rates of all species are very

low, but without necessarily a higher growth rate in P. aspleniifolius than

in co-occurring species. The results of the growth rate studies are

integrated with some other aspects of the biology of these species in a

discussion of the mechanisms determining canopy composition of Tasmanian

rain forests.

 

Skinner, M. F. and P. M. Attiwill (1981). "The Productivity of Pine

Plantations in Relation to Previous Land Use II. Phosphorus Adsorption

Isotherms and the Growth of Pine Seedlings." Plant And Soil 61(3): 329-339.

The improved growth of pine (Pinus radiata ) plantations on pasture soils

compared with that on soils which previously supported native eucalypt

forest is primarily explained in terms of soil phosphorus. Pasture

development has resulted in a decrease in the P adsorption maximum of about

300 mu g g super(-1) soil, a figure which agress with the increase in total

P due to the application of superphosphate. P adsorption isotherms were

used to calculate additions of P to give comparable levels of soil solution

P in eucalypt and pasture soils. The growth of pine seedlings in soils thus

amended showed a strong N x P interaction. When P was non-limiting,

addition of N raised productivity of the eucalypt soil above that of the

pasture soil. It is postulated that the different nature of the N x P

interaction in eucalypt and pasture soils results from differences in the

nitrogen cycle in the two soils.

 

Thomas, I. and J. B. Kirkpatrick (1996). "The roles of coastlines, people

and fire in the development of heathlands in northeast Tasmania." J.

Biogeogr. 23(5): 717-728.

Pollen analysis of a core taken from a reed marsh in northeastern Tasmania

near Bass Strait highlights the interplay between climatic changes, sea

level rises and the effects of Aborigines during the Holocene. Prior to

10,000 BP the region formed part of the Bassian landbridge which connected

Tasmania to the Australian mainland. Vegetation at that time was

characterized by shrubby grasslands with a very sparse tree cover. After

9000 BP, Eucalyptus forests prospered until about 6000 BP. A combination of

salt spray from a stabilizing mid-Holocene coast, the progressive effects

of soil podsolization and burning by Aborigines, eventually favoured the

replacement of grassy Eucalyptus forests by Allocasuarina with an

increasingly heathy understorey. The treeless heaths dominated by dwarfed

Allocasuarina which presently dominate the area are regarded as part of a

degradation sequence in which the above factors are implicated.

 

Tyler, P. A. (1992). "A lakeland from the Dreamtime. The Second Founders'

Lecture." Br. Phycol. J no. 4: pp.

The mountainous wilderness of Tasmania's World Heritage Area and

contiguous land is a district of lakes and rivers of immense beauty and

interest. A congruence of change in climate, relief, geology, soils and

vegetation divides the island into western and eastern provinces. A jagged,

western land of ancient rocks is mantled by peat-forming rainforest and

sedgeland, where creeks run, unenriched with minerals, to topaz, red-window

lakes. Eastwards lies a younger, flatter land, covered by sclerophyll

forests of Eucalyptus . Minerals from the soluble rocks give the lakes

distinctive chemistry compared with the brown dilute sea-water which drains

the western quartz. No peat extracts stain these eastern lakes and they lie

crystal clear with deep green windows. In this wilderness is a rich

diversity of rare microscopic organisms. Some, long forgotten, have been

rediscovered there. Others, new and novel, turn up with every cast of the

net. Among the richest sites are the coastal, fresh-water lagoons which the

Aboriginal inhabitants would have known intimately. Beside the Gordon River

are small lakes of very special interest. Periodically, they are topped up

with salt water from the estuary, keeping them meromictic, with brackish

water below and fresh water above. Such lakes and their unusual features

are uncommon in the world. They have social relevance. Because of their

meromictic condition, their sediments hold an especially fine-resolution

chronology of prehistoric climates and vegetational changes which shaped

Aboriginal fortunes to the times of European contact. Tasmania must stand

as one of the finest lake districts of the world. Perhaps nowhere else is

there such limnological richness and diversity in so small an encompass as

this island. Add to this its predominantly pristine nature, its uniqueness

and its beauty, and we have in all respects a World Heritage wetland

unsurpassed in this degraded world.

 

West, P. and G. Osler (1995). "Growth response to thinning and its relation

to site resources in Eucalyptus regnans." Canadian Journal of Forest

Research 25(1): 69-80.

The factors determining individual tree growth response are examined

during the 4 years following thinning in experiments in even-aged, 8- or

12-year-old regrowth Eucalyptus regnans forest at one site in Tasmania and

one site in Victoria. At the Tasmanian site, a vigorous understorey

dominated by a sedge developed after the thinning. At that site, light-use

efficiency by the trees was unaffected by thinning and the aboveground

biomass production by the trees in the thinned stand was substantially less

than that in the unthinned stand. At the site in Victoria, little

understorey developed, light-use efficiency by trees in the thinned stand

was greater than that in the unthinned stand, and aboveground biomass

production was unaffected by thinning even though the leaf weight of the

thinned stand was far less than that of the unthinned stand. Where the

understorey developed, it was concluded that it competed successfully with

the trees for water, thereby reducing production in the thinned stand when

compared with the unthinned stand. The individual tree growth response that

occurred in the thinned stand at that site appeared to be due solely to the

extra light available to individual trees following the canopy opening.

Where the understorey did not develop, it was concluded that individual

tree growth response was due not only to the extra light available to

individual trees but also to the increased availability of belowground

resources, most probably soil water. Application of a pre-existing stand

growth model suggested that at that site the tendency for increased growth

resulting from extra water availability in the thinned stand was just

balanced by decreased growth due to lower radiation absorption by the

reduced canopy, so that net production was unaffected by thinning.

 

Weston, C. J. and P. M. Attiwill (1990). "Effects of fire and harvesting on

nitrogen transformations and ionic mobility in soils of Eucalyptus regnans

forests of south-eastern Australia." Oecologia 83(1): 20-26.

Effects of fire and forest harvesting on inorganic-N in the soil, on net

N-mineralization, and on the leaching of NO super(-)@)d3-N and metallic

cations were measured in forests of Eucalyptus regnans) following a severe

wildfire in 1983. The concentrations of NO super(-)@)d3-N and metallic

cations in soil solution increased with increasing fire intensity.

Processes which limit the production and persistence of NO super(-)@)d3-N

in soil solution following disturbance will significantly reduce nutrient

losses or redistribution.

 

Weston, C. J. and P. M. Attiwill (1996). "Clearfelling and burning effects

on nitrogen mineralization and leaching in soils of old-age Eucalyptus

regnans forests." For. Ecol. Manage. 89: 1-3.

An argument against clearfelling and burning operations in forests is that

nutrient reserves may be diminished, leading to productivity decline over

successive rotations. Nitrogen is of primary concern as it is readily

volatilised and may be leached and thus the retention of nitrogen is a key

recovery process following perturbation. In this study we measured N

mineralization in situ and nitrogen concentrations in soil water from an

old-age Eucalyptus regnans forest (about 250 years old) and from

clearfelled forest in which treatment areas of unburnt ground, burnt ground

and burnt ground maintained free from regrowth were established. Total

inorganic N in the soil (0-5 cm) increased to a maximum of 168 mu g/g of

dry soil in clearfelled forest following burning, compared with 33 mu g/g

of dry soil in undisturbed forest. Increased total inorganic N in the soil

returned to a concentration equal to that in undisturbed forest most

rapidly in the clearfelled unburnt forest (6-9 months) and persisted for

the longest amount of time in the most severely disturbed site (clearfelled

burnt+herbicide-treated forest; 15-18 months). Net annual N mineralization

in undisturbed forest soils (0-5 cm) averaged 74.9 mu g/g of dry soil over

the two years of the study. In contrast, annual average N mineralization

was negative in two of the three clearfelled sites with 148.3 mu g/g of dry

soil of N immobilized in clearfelled and burnt forest. The concentration of

NO sub(3-) in soil water increased with increasing forest disturbance. Over

the first 260 days following clearfelling NO sub(3-)-N concentrations in

soil water at 10 cm depth averaged 2.6 mg/l in undisturbed forest, 8.5 mg/l

in clearfelled unburnt forest, 24.2 mg/l in clearfelled burnt forest, and

60.3 mg/l in clearfelled burnt+herbicide treated forest. Studies of net N

mineralization in situ, and of NO sub(3-) in soil water, support the

hypothesis that inorganic N was immobilized in all disturbed forests.

Immobilization of N by soil micro-organisms is alone not sufficient to

limit nitrification and NO sub(3-) leaching in disturbed E. regnans

forests. Rapid uptake of N by regrowing vegetation is essential in reducing

the availability of substrate for nitrification (NH sub(4+)) as well as in

reducing NO sub(3-) concentrations in soil water. Clearfelling of the E.

regnans catchment in this study did not significantly increase streamwater

NO sub(3-) concentrations and demonstrates the resilience of E. regnans

forests to leaching losses of N following destructive disturbance.

 

Wilkinson, G., M. Battaglia, et al. (1993). "Silvicultural use and effects

of fire." Technical Bulletin Native Forest Silviculture, Forestry

Commission, Tasmania No. 11, 60 pp.; 11 pp. of ref. PUBLISHER

INFORMATION(Forestry Commission): Tasmamia.

A discussion is presented in 3 parts on the effects of fire on the

silvicultural management of Tasmania's native forests which are mostly

eucalypt (Eucalyptus spp.) forests. Part A provides guidelines for the

silvicultural use of fire for site preparation and fuel reduction and for

the management of forests damaged by fire. Part B contains a summary of the

ecological effects of fire in Tasmania's forests. Part C is a comprehensive

literature review (on which parts A and B are based) and includes sections

on (i) prehistory of fire in Tasmania, (ii) eucalypt fire ecology, (iii)

ecological models of vegetation, fire and time, and (iv) effects of fire on

soils, water, air, vegetation and fauna.